The Murmillo is the main Atlantean infantry. They are strong against cavalry and weak against archers like the Toxotes and counter-infantry like the Axeman or Hypaspist.

Atlantean warriors and soldiers were known for their discipline and drill. A Murmillo began drill before dawn and finished at sunset. They were called Murmillo, or fish, because their shining, finned helmets made their complex movements in combat look like a school of swimming fish. They have a rivalry with and respect for the Destroyers, who consider the Murmillo beneath them - as a fisherman would a fish.